Reaction of rat liver glutathione S-transferases and bacterial dichloromethane dehalogenase with dihalomethanes

Blocki, FA; Logan, MS; Baoli, C; Wackett, LP

HERO ID

730517

Reference Type

Journal Article

Year

1994

Language

English

PMID

8132617

HERO ID 730517
In Press No
Year 1994
Title Reaction of rat liver glutathione S-transferases and bacterial dichloromethane dehalogenase with dihalomethanes
Authors Blocki, FA; Logan, MS; Baoli, C; Wackett, LP
Journal Journal of Biological Chemistry
Volume 269
Issue 12
Page Numbers 8826-8830
Abstract Dichloromethane dehalogenase from Methylophilus sp. DM11 is a glutathione S-transferase homolog that is specifically active with dihalomethane substrates. This bacterial enzyme and rat liver glutathione S-transferases were purified to investigate their relative reactivity with CH2Cl2 and related substrates. Rat liver alpha class glutathione transferases were inactive and mu class enzymes showed low activity (7-23 nmol/min/mg of protein) with CH2Cl2. theta class glutathione transferase 5-5 from rat liver and Methylophilus sp. dichloromethane dehalogenase showed specific activities of > or = 1 mumol/min/mg of protein. Apparent Kcat/Km were determined to be 3.3 x 10(4) and 6.0 x 10(4) L M-1 S-1 for the two enzymes, respectively. Dideutero-dichloromethane was processed to dideutereo-formaldehyde, consistent with a nucleophilic halide displacement mechanism. The possibility of a GSCH2X reaction intermediate (GS, glutathione; X, halide) was probed using CH2ClF to generate a more stable halomethylglutathione species (GSCH2F). The reaction of CH2ClF with dichloromethane dehalogenase produced a kinetically identifiable intermediate that decomposed to formaldehyde at a similar rate to synthetic HOCH2CH2SCH2F. 19F-NMR revealed the transient formation of an intermediate identified as GSCH2F by its chemical shift, its triplet resonance, and H-F coupling constant consistent with a fluoromethylthioether. Its decomposition was matched by a stoichiometric formation of fluoride. These studies indicated that the bacterial dichloromethane dehalogenase directs a nucleophilic attack of glutathione on CH2Cl2 to produce a halomethylthioether intermediate. This focuses attention on the mechanism used by theta class glutathione transferases to generate a halomethylthioeter from relatively unreactive dihalomethanes.
Pmid 8132617
Wosid WOS:A1994NB41100034
Url <Go to ISI>://WOS:A1994NB41100034
Is Certified Translation No
Dupe Override No
Is Public Yes
Language Text English
Is Qa No